1. Department of Physics & Astronomy, University College London, Gower Street, London WC1E 6BT, United Kingdom
2. Centre for Mathematics and Physics in the Life Sciences and Experimental Biology (CoMPLEX), University College London, Gower Street, London WC1E 6BT, United Kingdom
3. Research Department of Cell & Developmental Biology, University College London, Gower Street, London WC1E 6BT, United Kingdom