1. SUPA, School of Physics and Astronomy, University of St. Andrews, St. Andrews, Fife KY16 9SS, United Kingdom
2. Carnegie Physics Laboratory and Division of Molecular Medicine, University of Dundee, Dundee DD1 4HN, United Kingdom
3. School of Biology, Medical and Biological Sciences Building, North Haugh, University of St. Andrews, St. Andrews, Fife KY16 9TF, United Kingdom