1. School of Biology, College of Sciences, University of Tehran, Tehran, Iran
2. Department of Biology, Faculty of Sciences, Ferdowsi University of Mashhad, Mashhad, Iran. *
3. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan 685000, Russia
4. Department of Biology, Faculty of Sciences, University of Isfahan, Isfahan, Iran
5. Naturhistorisches Museum Basel, Augustinergasse, Switzerland