1. Zoological Museum, Biodiversity Unit, University of Turku, FI-20014, Finland email:
2. A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nakhimov Ave. 2, Sevastopol, 299011 email:
3. Perm State University, Bukireva Street 15, Perm, 614600, Russia email:
4. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia and Department of Zoology & Entomology, University of the Free State, Bloemfontein, 9300, South Africa email: