1. Division of Medical Sciences and Graduate Entry Medicine, University of Nottingham, Royal Derby Hospital, Derby, UK
2. Ninewells Hospital, Dundee, UK
3. Division of Epidemiology and Public Health, School of Medicine, University of Nottingham, Nottingham, UK
4. National Institute for Health Research Nottingham Digestive Diseases Biomedical Research Unit, Nottingham University Hospitals NHS Trust, Nottingham, UK