1. 1Department of Medical Sciences, School of Veterinary Medicine, University of Wisconsin–Madison, Madison, Wisconsin 53706
2. 2Department of Veterinary Clinical Medicine, School of Veterinary Medicine, University of Wisconsin–Madison, Madison, Wisconsin 53706
3. 3Department of Veterinary Pathobiology, College of Veterinary Medicine, University of Illinois at Urbana–Champaign, Urbana, Illinois 61802, USA