1. 1Laboratory of Marine Biology, Graduate School of Fisheries Science, Hokkaido University, 3-1-1 Minato-cho, Hakodate, Hokkaido, 041-8611, Japan
2. 2Arctic Environmental Research Centre, National Institute of Polar Research, 10-3 Midori-cho, Tachikawa, Tokyo, 190-8518, Japan
3. 3Hokkaido Research Organisation, Central Fisheries Research Institute, 238 Hamanaka-cho, Yoichi, Hokkaido, 046-8555, Japan