1. Graduate School of Biosphere Science, Hiroshima University, 1-4-4 Kagamiyama, Higashi-hiroshima, Hiroshima 739-8528, Japan
2. Graduate School of Marine Science and Technology, Tokyo University of Marine Science and Technology, 4-5-7 Konan, Minato, Tokyo 108-8477, Japan
3. Department of Environment and Agriculture, School of Science, Curtin University, Technology Park, 1 Turner Avenue, Bentley, WA 6102, Australia