1. Department of Infection Biology; School of Veterinary Science; Institute of infection and Global Health; Liverpool Merseyside L3 5RF UK
2. University of Liverpool & Wood Veterinary Group; Gloucester GL2 4NB UK
3. Department of Epidemiology and Population Health; Institute of Infection and Global Health; University of Liverpool; Leahurst Campus; Neston Wirral CH64 7TE UK