1. Department of Epidemiology and Population Health; Institute of Infection and Global Health, University of Liverpool; Liverpool Science Park IC2, 146 Brownlow Hill Liverpool L3 5RF UK
2. Department of Infection Biology; Institute of Infection and Global Health, School of Veterinary Science, University of Liverpool; Liverpool Science Park IC2, 146 Brownlow Hill Liverpool L3 5RF UK
3. University of Liverpool & Wood Veterinary Group; Gloucester Gloucestershire GL2 4NB UK