1. The Philip Leverhulme Equine Hospital; School of Veterinary Science; University of Liverpool; Neston Wirral CH64 7TE UK
2. The Philip Leverhulme Equine Hospital; School of Veterinary Science; The University of Liverpool; Leahurst Campus Neston Wirral CH64 7TE UK
3. Department of Obesity and Endocrinology; Institute of Ageing and Chronic Disease; The University of Liverpool; Leahurst Campus Neston Wirral CH64 7TE UK
4. The Queen's Medical Research Institute; University of Edinburgh; 47 Little France Cresent Edinburgh Scotland EH16 4TJ UK
5. Department of Musculoskeletal Biology; Institute of Ageing and Chronic Disease; The University of Liverpool; Leahurst Campus Neston Wirral CH64 7TE UK