1. CSIRO Plant Industry, North Ryde, NSW 1670, Australia. Present address for Gianibelli and Larroque: CSIRO Plant Industry, GPO Box 1600, Canberra, ACT 2601, Australia.
2. Corresponding author. E-mail: giani@pi.csiro.au
3. Quality Wheat CRC, North Ryde, NSW 1670, Australia.
4. Grain Science and Industry, Kansas State University, Manhattan, KS 66506-2201.