1. Department of Ocean Sciences Tokyo University of Marine Science and Technology Tokyo Japan
2. Institute for Marine and Antarctic Studies University of Tasmania Hobart Tasmania Australia
3. Atmosphere and Ocean Research Institute The University of Tokyo Chiba Japan
4. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences The University of Tokyo Tokyo Japan
5. Japan Fisheries Research and Education Agency Fisheries Resources Institute Kawasaki Japan
6. Japan Fisheries Research and Education Agency Fisheries Resources Institute Nagasaki Japan
7. Japan Fisheries Research and Education Agency Fisheries Technology Institute Okinawa Japan
8. Graduate School of Science and Engineering Soka University Tokyo Japan