1. Department of Neuroscience, University of Wisconsin Medical School, Madison, Wisconsin;
2. INSERM U587, Université Victor Segalen Bordeaux 2, Hôpital Pellegrin, Bordeaux, France;
3. Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom; and
4. Institute for Science and Technology in Medicine, School of Life Sciences, Keele University, Keele, United Kingdom