1. Laboratory of Plants Improvement and Valorization of Agroressources (LAPVA), National School of Engineering of Sfax (ENIS), University of Sfax, 3038 Sfax, Tunisia
2. University of Sfax, High Institute of Biotechnology, Sfax, Tunisia
3. University of Sfax, Faculty of Science, LMSE, BP 802, 3018 Sfax, Tunisia
4. Department of Life Sciences, Faculty of Sciences of Gafsa, Gafsa University, 2112 Gafsa, Tunisia
5. Univ. Grenoble Alpes, CNRS, CERMAV, F-38000 Grenoble, France