1. Centre de Recherche en Biologie Forestière, Pavillon C‐E‐Marchand, Université Laval, Québec G1K 7P4, Canada
2. To whom correspondence should be addressed at: Eastern Regional Research Center, ARS/USDA, 600 East Mermaid Lane, Wyndmoor, PA 19038, USA. E‐mail:
3. Present address: Eastern Regional Research Center, ARS/USDA, 600 East Mermaid Lane, Wyndmoor, PA 19038, USA. E‐mail:
4. Department of Biology and Microbiology, University of Wisconsin Oshkosh, 800 Algoma Bvd., Oshkosh, WI 54901–8640, USA
5. International Institute of Biotechnology, Department of Biosciences, University of Kent, Canterbury, Kent CT2 7YW, UK
6. Recherches en Sciences de la Vie et de la Santé, Pavillon C‐E‐Marchand, Université Laval, Québec G1K 7P4, Canada