1. U.S. Forest Service, Institute of Pacific Islands Forestry, 23 E. Kawili Street, Hilo, HI 96720, U.S.A.,
email
2. Department of Ecology and Evolutionary Biology, University of California, Irvine, CA 92717, U.S.A.
3. National Tropical Botanical Garden, Lawai, HI 96765, U.S.A.
4. Department of Biological Sciences, Stanford University, Stanford, CA 94305–5020, U.S.A.