1. Department of Plant Genetics and Biotechnology, Horticulture Research International, Wellesbourne, Warwick, CV35 9EF, UK,
2. Department of Crop Physiology, Horticulture Research International, East Malling, West Malling, Kent, ME19 6BJ, UK and
3. Massey University, Institute of Food, Nutrition & Human Health, Palmerston North, New Zealand