1. Institute of Biological Sciences, University of Tsukuba, Tsukuba, Ibaraki, 305‐8572 Japan,
2. Global Environment Research Group, National Institute for Environmental Studies, Tsukuba, Ibaraki, 305‐0053 Japan,
3. Department of Biology, Graduate School of Science, Osaka University, 1‐16, Machikaneyama‐cho, Toyonaka, Osaka, 560‐0043 Japan, and
4. Institute for Basin Ecosystem Studies, Gifu University, Yanagido, Gifu, 501‐1193, Japan