1. Graduate School of Pharmaceutical Sciences, Kyoto University, Sakyo-ku, Kyoto 606-8501, Japan
2. Graduate School of Medicine, Kyoto University, Sakyo-ku, Kyoto 606-8507, Japan
3. RIKEN Center for Computational Science, Chuo-ku, Kobe 650-0047, Japan
4. Research Institute for Microbial Diseases, Osaka University, Suita 565-0871, Japan
5. Immunology Frontier Research Center (IFReC), Osaka University, Suita 565-0871, Japan