1. National Research Institute of Brewing, 3-7-1 Kagamiyama, Higashihiroshima 739-0046, Japan; Department of Biofunctional Science and Technology, Graduate School of Biosphere Science, Hiroshima University, 1-4-4 Kagamiyama, Higashihiroshima 739-8528, Japan; and Department of Chemistry, Graduate School of Science, Hiroshima University, Higashihiroshima 739-8528, Japan