1. Shandong Key Laboratory of Environmental Processes and Health, School of Environmental Science and Engineering, Shandong University, Qingdao, Shandong 266237, P. R. China
2. State Key Laboratory of Microbial Technology, Shandong University, Qingdao 266237, P. R. China
3. Sino-French Research Institute for Ecology and Environment (ISFREE), Shandong University, Qingdao, Shandong 266237, P. R. China
4. College of Geography and Environment, Shandong Normal University, Jinan, Shandong 250014, P. R. China