1. School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth, PL4 8AA, United Kingdom
2. School of Geography, Earth and Environmental Sciences, University of Plymouth, Drake Circus, Plymouth, PL4 8AA, United Kingdom
3. Charles River, Elpinstone Research Centre, Elphinstone, Tranent EH33 2NE, United Kingdom
4. Institut des Sciences de la Mer de Rimouski (ISMER), Université du Québec à Rimouski, 310 allée des Ursulines, Rimouski, Québec Canada G5L 3A1
5. Institute of Life and Earth Sciences Heriot-Watt University, John Muir Building, Edinburgh, EH14 4AS, United Kingdom