1. Marine Environmental Sciences, Fisheries and Oceans Canada, Bedford Institute of Oceanography, Dartmouth, Nova Scotia B2Y 4A2, Canada, 343 East 11th Street, North Vancouver, British Columbia V7L 2H1, Canada, 31 Brentlawn Boulevard, Winnipeg, Manitoba R3T 4X9, Canada, and Fisheries and Oceans Canada, Central and Arctic Region, Freshwater Institute, 501 University Crescent, Winnipeg, Manitoba R3T 2N6, Canada