1. CAS Key Laboratory of Biofuels and Shandong Provincial Key Laboratory of Synthetic Biology, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, Shandong 266101, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Shandong Energy Institute, Qingdao, Shandong 266101, China
4. Qingdao New Energy Shandong Laboratory, Qingdao, Shandong 266101, China