1. Osaka University, 1-3 Machikaneyama-cho, Toyonaka, Osaka 560-8531, Japan
2. Department of Fundamental Bioscience, Shiga University of Medical Science, Otsu, Shiga 520-2192, Japan
3. Advanced Research Laboratories, Tokyo City University, 8-15-1 Todoroki, Setagaya, Tokyo 158-0082, Japan