1. Bioresources Research and Development Department, Sapporo Breweries Ltd., 37-1, Nittakizaki, Ota, Gunma 370-0393, Japan, Frontier Laboratories of Value Creation, Sapporo Breweries Ltd., 10 Okatohme, Yaizu, Shizuoka, 425-0013, Japan, and Barley Germplasm Center, Research Institute for Bioresources, Okayama University, 2-20-1, Chuo, Kurashiki, 710-0046, Japan