1. Key Laboratory of Unconventional Oil & Gas Development, Ministry of Education, School of Petroleum Engineering, China University of Petroleum (East China), Qingdao, Shandong 266580, China
2. Baikouquan Oil Production Plant of PetroChina Xinjiang Oilfield Branch, Kelamayi, Xinjiang 834000, China
3. Centre for Bioengineering and Biotechnology, China University of Petroleum (East China), Qingdao, Shandong 266580, China