1. Plant Biotechnology Department, National Institute of Agrobiological Sciences, 2-1-2 Kannondai, Tsukuba, Ibaraki 305-8602, Japan; Faculty of Life Science and Biotechnology, Fukuyama University, Gakuencho 1, Fukuyama, Hiroshima 729-0292, Japan; and Agriculture, Forestry and Fisheries Research Council Secretariat, 1-2-1 Kasumigaseki Chiyodaku, Tokyo 100-8950, Japan