1. Department of Biology, University of Ottawa, 30 Marie Curie Street, Ottawa, Ontario, Canada K1N 6N5; Department of Biochemistry, Microbiology and Immunology, University of Ottawa, 451 Smyth Road, Ottawa, Ontario, Canada K1H 8M5; and Eastern Cereal and Oilseed Research Center, Agriculture and Agri-Food Canada, Ottawa, Ontario, Canada K1A 0C6