1. Institute of Ageing and Chronic Disease, University of Liverpool, Liverpool L7 8TX, U.K.
2. Institute of Integrative Biology, University of Liverpool, Liverpool L69 7ZB, U.K.
3. HLS Technology Directorate, University of Liverpool, Liverpool L69 3GB, U.K.
4. Institute of Veterinary Science, University of Liverpool, Leahurst Campus, Neson CH64 7TE, U.K.