1. Department of Biotechnology, Faculty of Technology, Tokyo University of Agriculture and Technology, Tokyo 184-8588, Japan, Center for Food and Environmental Sciences, The Tokyo Kembikyoin Foundation, 5-3-11, Maesawa, Higashi-kurume, Tokyo 203-0032, Japan, and Department of Applied and Biological Science, Faculty of Agriculture, Tokyo University of Agriculture and Technology, Tokyo 183-8509, Japan