1. Department of Economic and Food Sciences, Section of Food Technology and Biotechnology, University of Perugia, Via S. Costanzo n.c.n., 06126 Perugia, Italy
2. Department of Applied Biology, Section of Microbiology, University of Perugia, Via Borgo XX Giugno 74, 06121 Perugia, Italy
3. Italian Brewing Research Center (Centro di Eccellenza per la Ricerca sulla Birra—CERB), University of Perugia, Via S. Costanzo n.c.n., 06126 Perugia, Italy