1. Plants, Soils, and Climate, College of Agriculture and Applied Sciences (CAAS), Utah State University (USU), Logan, Utah 84322, United States
2. U.S. Salinity Laboratory (USDA-ARS), 450 W Big Springs Road, Riverside, California 92507, United States
3. College of Natural and Agricultural Sciences, University of California Riverside, 900 University Avenue, Riverside, California 92521, United States
4. Department of Botany & Plant Sciences, University of California Riverside, Riverside, California 92521, United States