1. Department of Agrobioscience, Graduate School of Agriculture, Kobe University, Nada, Kobe 657-8501, Japan
2. Department of Molecular Microbiology, Faculty of Life Sciences, Tokyo University of Agriculture, Sakuragaoka, Setagaya, Tokyo 156-8502, Japan
3. Faculty of Science, Ibaraki University, Mito 310-8512, Japan
4. Institute for Extra-Cutting-Edge Science and Technology Avant-garde Research (X-star), Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Kanagawa 237-0061, Japan
5. Department of Microbiology, Southern Illinois University, Carbondale, Illinois 62901, United States