1. Institute for Bee Products & Health Science, Yamada Bee Company, Inc., 194 Ichiba, Kagamino, Okayama 708-0393, Japan
2. RIKEN Center for Sustainable Resource Science, 2-1 Wako, Saitama 351-0198, Japan
3. Graduate School of Advanced Life Science, Frontier Research Center for Post-Genome Science and Technology, Hokkaido University, N21W11, Sapporo 001-0021, Japan