1. National Food Research Institute, 2-1-12 Kannondai, Tsukuba, Ibaraki 305-8642, Japan, Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1, Tennoudai, Tsukuba, Ibaraki 305-8577, Japan, and Japan Society for the Promotion of Science, 6 Ichibancho, Chiyoda-ku, Tokyo 102-8471, Japan