1. Department of Wildlife, Fish & Conservation Biology, University of California, Davis, Davis, California 95616, United States
2. School of Veterinary Medicine, Department of Anatomy, Physiology and Cell Biology, University of California, Davis, California 95616, United States
3. U.S. Geological Survey, California Water Science Center, 6000 J Street, Placer Hall, Sacramento, California 95819, United States
4. Department of Civil and Environmental Engineering, University of California, Davis, California 95616, United States
5. Fish Conservation and Culture Laboratory, Department of Biological and Agricultural Engineering, University of California, 17501 Byron Hwy, Byron, Davis, California 94514, United States