1. Marine Ecology and Biodiversity, Plymouth Marine Laboratory, Prospect Place, West Hoe, Plymouth PL1 3DH, U.K.
2. School of Life Sciences, University of Essex, Wivenhoe Park, Colchester CO4 3SQ, U.K.
3. Plymouth Marine Laboratory, Prospect Place, West Hoe, Plymouth PL1 3DH, U.K.
4. Marine Biogeochemistry and Ocean Observations, Plymouth Marine Laboratory, Prospect Place, West Hoe, Plymouth PL1 3DH, U.K.
5. Marine Biology and Ecology Research Centre (MBERC), School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL4 8AA, U.K.