1. Stockbridge School of Agriculture and School of Earth & Sustainability, University of Massachusetts, Amherst, Massachusetts 01003, United States
2. Department of Biology, Lund University, 223 62 Lund, Sweden
3. Department of Geosciences, University of Massachusetts, Amherst, Massachusetts 01003, United States
4. Department of Environmental Science, University of Arizona, Tucson, Arizona 85719, United States
5. The Ecosystems Center, Marine Biological Laboratory, Woods Hole, Massachusetts 02543, United States