1. Department of Urban Engineering, School of Engineering, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-8654, Japan
2. Center for Marine Environmental Studies (CMES), Ehime University, Bunkyo-cho 3, Matsuyama-shi, Ehime 790-8577, Japan
3. Department of Virology, Toyama Institute of Health, 17-1 Nakataikoyama, Imizu-shi, Toyama 939-0363, Japan
4. Laboratory of Viral Infection I, Department of Infection Control and Immunology, O̅mura Satoshi Memorial Institute & Graduate School of Infection Control Sciences, Kitasato University, Tokyo 108-8641, Japan