1. Center for Mitochondrial and Epigenomic Medicine, Children’s Hospital of Philadelphia, Philadelphia, Pennsylvania 19104, United States
2. Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, St. Louis, Missouri 63110, United States
3. Donald Danforth Plant Science Center, St. Louis, Missouri 63132, United States
4. Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania 19104, United States