1. Current address: USDA-ARS Hard Winter Wheat Genetics Research Unit, 4008 Throckmorton Hall, Kansas State University, Manhattan, KS 66506, U.S.A.
2. Current address: Dept. of Crop Science, 840 Main Campus Dr., Campus Box 7287, North Carolina State University, Raleigh, NC 27513, U.S.A.
3. Current address: VTT Biotechnology and Research, P.O. Box 1501 (Tietotie 2) FIN-02044 VTT, Finland
4. Address: USDA-ARS Cereal Crops Research Unit, 502 Walnut St., Madison, WI 53726, U.S.A.