1. State Key Laboratory of Microbial Technology and Marine Biotechnology Research Center, Shandong University, 72 Binhai Road, Qingdao 266237, Shandong, P. R. China
2. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology, 168 Wenhai Road, Qingdao 266237, Shandong, P. R. China