1. Department of Chemistry and Biomolecular Sciences, University of Ottawa, 10 Marie Curie, Ottawa, Ontario K1N 6N5, Canada
2. Biophysics Graduate Group, University of California, Berkeley, Berkeley, California 94720, United States
3. Department of Physics, University of California, Berkeley, Berkeley, California 94720, United States
4. Department of Molecular and Cell Biology, University of California, Berkeley, Berkeley, California 94720, United States
5. Institute for Quantitative Biosciences-QB3, University of California, Berkeley, Berkeley, California 94720, United States