1. Arthropod Genetics, The Pirbright Institute, Ash Road, Pirbright GU24 0NF, U.K.
2. Department of Zoology, University of Oxford, 11a Mansfield Road, Oxford OX1 3SZ, U.K.
3. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk NR4 7TJ, U.K.