1. Komaba Open Laboratory (KOL) and Research Center for Advanced Science and Technology (RCAST), The University of Tokyo, 4-6-1 Komaba, Meguro-ku, Tokyo 153-8904, Japan, Graduate School of Science and Technology, Nihon University, 1-8-14 Surugadai, Kanda, Chiyoda-ku, Tokyo 101-8308, Japan, and Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), 4-1-8 Honcho, Kawaguchi, Saitama 332-0012, Japan