1. Department of Basic Sciences, Graduate School of Arts and Sciences, The University of Tokyo, 3-8-1 Komaba, Meguro-ku, Tokyo 153-8902 Japan
2. Research Center for Complex Systems Biology, The University of Tokyo, 3-8-1 Komaba, Meguro-ku Tokyo 153-8902 Japan
3. PRESTO, Japan Science and Technology Agency (JST), 4-1-8 Honcho Kawaguchi, Saitama 332-0012, Japan