1. Watts & Crane Associates, 23 London Street, Faringdon, Oxfordshire, SN7 7AG, Department of Ecology & Biodiversity, The Swire Institute of Marine Science, The University of Hong Kong, Pokfulam, Hong Kong, P.R. China, Environment Agency Science Group, Evenlode House, Howbery Park, Wallingford, Oxfordshire, OX10 8BD, and Department of Statistics and Actuarial Science, The University of Hong Kong, Pokfulam, Hong Kong, P.R. China