1. Department of Synthetic Chemistry and Biological Chemistry, Graduate School of Engineering, Kyoto University, PRESTO, Japan Science and Technology Corporation (JST), Sakyo-ku, Yoshida Hommachi, Kyoto 606-8501, Japan, Section of Physiological Anatomy, Fukui Prefectural University, Faculty of Nursing and Welfare, 4-1-1 Kenjojima, Matsuoka-cho, Yoshida-gun, Fukui, 910-1195, Japan